The membranous vesicle of a submitochondrial particle after removal of the F1 spheres would be able to (1) oxidize NADH; (2) produce H2O from O2; (3) generate a proton gradient; (4) phosphorylate ADP. Which of the previous statements are true? How would this answer be different, if at all, if the objects being studied were intact submitochondrial particles that had been treated with dinitrophenol?